diff --git a/font-test-data/test_data/colrv1_json/colrv1_clipbox b/font-test-data/test_data/colrv1_json/colrv1_clipbox
index 1a512531b..1f382967e 100644
--- a/font-test-data/test_data/colrv1_json/colrv1_clipbox
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diff --git a/font-test-data/test_data/colrv1_json/colrv1_clipbox_CLIO_200 b/font-test-data/test_data/colrv1_json/colrv1_clipbox_CLIO_200
index 2d048af2c..f4b126a58 100644
--- a/font-test-data/test_data/colrv1_json/colrv1_clipbox_CLIO_200
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index 46eb50234..d24d42b9e 100644
--- a/font-test-data/test_data/colrv1_json/colrv1_colored_circles_v0
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diff --git a/font-test-data/test_data/colrv1_json/colrv1_composite_mode b/font-test-data/test_data/colrv1_json/colrv1_composite_mode
index 8d63e6fa1..91634d59e 100644
--- a/font-test-data/test_data/colrv1_json/colrv1_composite_mode
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--- a/font-test-data/test_data/colrv1_json/colrv1_paint_rotate
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index 9a44742b8..fd17e0ffc 100644
--- a/font-test-data/test_data/colrv1_json/colrv1_paint_rotate_ROTA_40
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index 9e9ac7fb3..9e007ae8a 100644
--- a/font-test-data/test_data/colrv1_json/colrv1_paint_rotate_ROTX_-250_ROTY_-250
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diff --git a/font-test-data/test_data/colrv1_json/colrv1_paint_scale b/font-test-data/test_data/colrv1_json/colrv1_paint_scale
index a820b0c8b..2baeb174f 100644
--- a/font-test-data/test_data/colrv1_json/colrv1_paint_scale
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diff --git a/font-test-data/test_data/colrv1_json/colrv1_paint_scale_SCOX_200_SCOY_200 b/font-test-data/test_data/colrv1_json/colrv1_paint_scale_SCOX_200_SCOY_200
index 31afc04ff..6ce279549 100644
--- a/font-test-data/test_data/colrv1_json/colrv1_paint_scale_SCOX_200_SCOY_200
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diff --git a/font-test-data/test_data/colrv1_json/colrv1_paint_scale_SCSX_-1_SCOY_-1 b/font-test-data/test_data/colrv1_json/colrv1_paint_scale_SCSX_-1_SCOY_-1
index 6ffc79177..011bdee8c 100644
--- a/font-test-data/test_data/colrv1_json/colrv1_paint_scale_SCSX_-1_SCOY_-1
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diff --git a/font-test-data/test_data/colrv1_json/colrv1_paint_scale_SCSX_0.25_SCOY_0.25 b/font-test-data/test_data/colrv1_json/colrv1_paint_scale_SCSX_0.25_SCOY_0.25
index 74dc6e1c4..512c73c1b 100644
--- a/font-test-data/test_data/colrv1_json/colrv1_paint_scale_SCSX_0.25_SCOY_0.25
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diff --git a/font-test-data/test_data/colrv1_json/colrv1_paint_skew b/font-test-data/test_data/colrv1_json/colrv1_paint_skew
index 17a85443a..d5ab2c081 100644
--- a/font-test-data/test_data/colrv1_json/colrv1_paint_skew
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--- a/font-test-data/test_data/colrv1_json/colrv1_paint_skew_SKCX_200_SKCY_200
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--- a/font-test-data/test_data/colrv1_json/colrv1_paint_skew_SKXA_20
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--- a/font-test-data/test_data/colrv1_json/colrv1_paint_skew_SKYA_20
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index 7f134ac85..d05709518 100644
--- a/font-test-data/test_data/colrv1_json/colrv1_paint_transform
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index 1f613ed0c..b6f3ce6d4 100644
--- a/font-test-data/test_data/colrv1_json/colrv1_paint_translate
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--- a/font-test-data/test_data/colrv1_json/colrv1_paint_translate_TLDX_100_TLDY_100
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+{"glyph_id":76,"ops":[{"PushClipBox":{"clip_box":{"x_min":0.0,"y_min":0.0,"x_max":1000.0,"y_max":1000.0}}},{"FillGlyph":{"gid":176,"transform":{"xx":1.0,"yx":0.0,"xy":0.0,"yy":1.0,"dx":0.0,"dy":0.0},"brush":{"SweepGradient":{"c0":{"x":500.0,"y":600.0},"start_angle":213.75,"end_angle":281.25,"color_stops":[{"offset":0.0,"palette_index":7,"alpha":1.0},{"offset":0.5,"palette_index":4,"alpha":1.0},{"offset":0.8333334,"palette_index":0,"alpha":1.0},{"offset":1.0,"palette_index":8,"alpha":1.0}],"extend":"Repeat"}}}},"PopClip"]}
+{"glyph_id":77,"ops":[{"PushClipBox":{"clip_box":{"x_min":0.0,"y_min":0.0,"x_max":1000.0,"y_max":1000.0}}},{"FillGlyph":{"gid":176,"transform":{"xx":1.0,"yx":0.0,"xy":0.0,"yy":1.0,"dx":0.0,"dy":0.0},"brush":{"SweepGradient":{"c0":{"x":500.0,"y":600.0},"start_angle":146.25,"end_angle":348.75,"color_stops":[{"offset":0.0,"palette_index":7,"alpha":1.0},{"offset":0.5,"palette_index":4,"alpha":1.0},{"offset":0.8333334,"palette_index":0,"alpha":1.0},{"offset":1.0,"palette_index":8,"alpha":1.0}],"extend":"Repeat"}}}},"PopClip"]}
+{"glyph_id":78,"ops":[{"PushClipBox":{"clip_box":{"x_min":0.0,"y_min":0.0,"x_max":1000.0,"y_max":1000.0}}},{"FillGlyph":{"gid":176,"transform":{"xx":1.0,"yx":0.0,"xy":0.0,"yy":1.0,"dx":0.0,"dy":0.0},"brush":{"SweepGradient":{"c0":{"x":500.0,"y":600.0},"start_angle":11.25,"end_angle":78.75,"color_stops":[{"offset":0.0,"palette_index":7,"alpha":1.0},{"offset":0.5,"palette_index":4,"alpha":1.0},{"offset":0.8333334,"palette_index":0,"alpha":1.0},{"offset":1.0,"palette_index":8,"alpha":1.0}],"extend":"Repeat"}}}},"PopClip"]}
+{"glyph_id":79,"ops":[{"PushClipBox":{"clip_box":{"x_min":0.0,"y_min":0.0,"x_max":1000.0,"y_max":1000.0}}},{"FillGlyph":{"gid":176,"transform":{"xx":1.0,"yx":0.0,"xy":0.0,"yy":1.0,"dx":0.0,"dy":0.0},"brush":{"SweepGradient":{"c0":{"x":500.0,"y":600.0},"start_angle":315.0,"end_angle":315.0,"color_stops":[{"offset":0.0,"palette_index":8,"alpha":1.0},{"offset":0.16666663,"palette_index":0,"alpha":1.0},{"offset":0.5,"palette_index":4,"alpha":1.0},{"offset":1.0,"palette_index":7,"alpha":1.0}],"extend":"Repeat"}}}},"PopClip"]}
+{"glyph_id":80,"ops":[{"PushClipBox":{"clip_box":{"x_min":0.0,"y_min":0.0,"x_max":1000.0,"y_max":1000.0}}},{"FillGlyph":{"gid":176,"transform":{"xx":1.0,"yx":0.0,"xy":0.0,"yy":1.0,"dx":0.0,"dy":0.0},"brush":{"SweepGradient":{"c0":{"x":500.0,"y":600.0},"start_angle":180.0,"end_angle":450.0,"color_stops":[{"offset":0.0,"palette_index":7,"alpha":1.0},{"offset":0.5,"palette_index":4,"alpha":1.0},{"offset":0.8333334,"palette_index":0,"alpha":1.0},{"offset":1.0,"palette_index":8,"alpha":1.0}],"extend":"Repeat"}}}},"PopClip"]}
+{"glyph_id":81,"ops":[{"PushClipBox":{"clip_box":{"x_min":0.0,"y_min":0.0,"x_max":1000.0,"y_max":1000.0}}},{"FillGlyph":{"gid":176,"transform":{"xx":1.0,"yx":0.0,"xy":0.0,"yy":1.0,"dx":0.0,"dy":0.0},"brush":{"SweepGradient":{"c0":{"x":500.0,"y":600.0},"start_angle":-100.00305,"end_angle":20.00061,"color_stops":[{"offset":0.0,"palette_index":8,"alpha":1.0},{"offset":0.16666663,"palette_index":0,"alpha":1.0},{"offset":0.5,"palette_index":4,"alpha":1.0},{"offset":1.0,"palette_index":7,"alpha":1.0}],"extend":"Repeat"}}}},"PopClip"]}
+{"glyph_id":82,"ops":[{"PushClipBox":{"clip_box":{"x_min":0.0,"y_min":0.0,"x_max":1000.0,"y_max":1000.0}}},{"FillGlyph":{"gid":176,"transform":{"xx":1.0,"yx":0.0,"xy":0.0,"yy":1.0,"dx":0.0,"dy":0.0},"brush":{"SweepGradient":{"c0":{"x":500.0,"y":600.0},"start_angle":-235.00305,"end_angle":155.00061,"color_stops":[{"offset":0.0,"palette_index":7,"alpha":1.0},{"offset":0.5,"palette_index":4,"alpha":1.0},{"offset":0.8333334,"palette_index":0,"alpha":1.0},{"offset":1.0,"palette_index":8,"alpha":1.0}],"extend":"Repeat"}}}},"PopClip"]}
+{"glyph_id":83,"ops":[{"PushClipBox":{"clip_box":{"x_min":0.0,"y_min":0.0,"x_max":1000.0,"y_max":1000.0}}},{"FillGlyph":{"gid":176,"transform":{"xx":1.0,"yx":0.0,"xy":0.0,"yy":1.0,"dx":0.0,"dy":0.0},"brush":{"SweepGradient":{"c0":{"x":500.0,"y":600.0},"start_angle":-337.48627,"end_angle":607.49725,"color_stops":[{"offset":0.0,"palette_index":8,"alpha":1.0},{"offset":0.16666663,"palette_index":0,"alpha":1.0},{"offset":0.5,"palette_index":4,"alpha":1.0},{"offset":1.0,"palette_index":7,"alpha":1.0}],"extend":"Repeat"}}}},"PopClip"]}
diff --git a/font-test-data/test_data/colrv1_json/colrv1_variable_alpha b/font-test-data/test_data/colrv1_json/colrv1_variable_alpha
index 9df9f0622..f396e4be2 100644
--- a/font-test-data/test_data/colrv1_json/colrv1_variable_alpha
+++ b/font-test-data/test_data/colrv1_json/colrv1_variable_alpha
@@ -1 +1 @@
-{"glyph_id":177,"ops":[{"PushClipBox":{"clip_box":{"x_min":0.0,"y_min":0.0,"x_max":1000.0,"y_max":1000.0}}},{"PushTransform":{"transform":{"xx":1.0,"yx":0.0,"xy":0.0,"yy":1.0,"dx":150.0,"dy":0.0}}},{"PushClipGlyph":{"gid":176}},{"FillBrush":{"brush":{"Solid":{"palette_index":3,"alpha":1.0}}}},"PopClip","PopTransform",{"PushTransform":{"transform":{"xx":1.0,"yx":0.0,"xy":0.0,"yy":1.0,"dx":-150.0,"dy":0.0}}},{"PushClipGlyph":{"gid":176}},{"FillBrush":{"brush":{"LinearGradient":{"p0":{"x":500.0,"y":250.0},"p1":{"x":500.0,"y":950.0},"color_stops":[{"offset":0.0,"palette_index":0,"alpha":1.0},{"offset":1.0,"palette_index":4,"alpha":1.0}],"extend":"Repeat"}}}},"PopClip","PopTransform","PopClip"]}
+{"glyph_id":177,"ops":[{"PushClipBox":{"clip_box":{"x_min":0.0,"y_min":0.0,"x_max":1000.0,"y_max":1000.0}}},{"PushTransform":{"transform":{"xx":1.0,"yx":0.0,"xy":0.0,"yy":1.0,"dx":150.0,"dy":0.0}}},{"FillGlyph":{"gid":176,"transform":{"xx":1.0,"yx":0.0,"xy":0.0,"yy":1.0,"dx":0.0,"dy":0.0},"brush":{"Solid":{"palette_index":3,"alpha":1.0}}}},"PopTransform",{"PushTransform":{"transform":{"xx":1.0,"yx":0.0,"xy":0.0,"yy":1.0,"dx":-150.0,"dy":0.0}}},{"FillGlyph":{"gid":176,"transform":{"xx":1.0,"yx":0.0,"xy":0.0,"yy":1.0,"dx":0.0,"dy":0.0},"brush":{"LinearGradient":{"p0":{"x":500.0,"y":250.0},"p1":{"x":500.0,"y":950.0},"color_stops":[{"offset":0.0,"palette_index":0,"alpha":1.0},{"offset":1.0,"palette_index":4,"alpha":1.0}],"extend":"Repeat"}}}},"PopTransform","PopClip"]}
diff --git a/font-test-data/test_data/colrv1_json/colrv1_variable_alpha_APH1_-0.7 b/font-test-data/test_data/colrv1_json/colrv1_variable_alpha_APH1_-0.7
index f2b615beb..21e8f5cdb 100644
--- a/font-test-data/test_data/colrv1_json/colrv1_variable_alpha_APH1_-0.7
+++ b/font-test-data/test_data/colrv1_json/colrv1_variable_alpha_APH1_-0.7
@@ -1 +1 @@
-{"glyph_id":177,"ops":[{"PushClipBox":{"clip_box":{"x_min":0.0,"y_min":0.0,"x_max":1000.0,"y_max":1000.0}}},{"PushTransform":{"transform":{"xx":1.0,"yx":0.0,"xy":0.0,"yy":1.0,"dx":150.0,"dy":0.0}}},{"PushClipGlyph":{"gid":176}},{"FillBrush":{"brush":{"Solid":{"palette_index":3,"alpha":0.2999878}}}},"PopClip","PopTransform",{"PushTransform":{"transform":{"xx":1.0,"yx":0.0,"xy":0.0,"yy":1.0,"dx":-150.0,"dy":0.0}}},{"PushClipGlyph":{"gid":176}},{"FillBrush":{"brush":{"LinearGradient":{"p0":{"x":500.0,"y":250.0},"p1":{"x":500.0,"y":950.0},"color_stops":[{"offset":0.0,"palette_index":0,"alpha":1.0},{"offset":1.0,"palette_index":4,"alpha":1.0}],"extend":"Repeat"}}}},"PopClip","PopTransform","PopClip"]}
+{"glyph_id":177,"ops":[{"PushClipBox":{"clip_box":{"x_min":0.0,"y_min":0.0,"x_max":1000.0,"y_max":1000.0}}},{"PushTransform":{"transform":{"xx":1.0,"yx":0.0,"xy":0.0,"yy":1.0,"dx":150.0,"dy":0.0}}},{"FillGlyph":{"gid":176,"transform":{"xx":1.0,"yx":0.0,"xy":0.0,"yy":1.0,"dx":0.0,"dy":0.0},"brush":{"Solid":{"palette_index":3,"alpha":0.2999878}}}},"PopTransform",{"PushTransform":{"transform":{"xx":1.0,"yx":0.0,"xy":0.0,"yy":1.0,"dx":-150.0,"dy":0.0}}},{"FillGlyph":{"gid":176,"transform":{"xx":1.0,"yx":0.0,"xy":0.0,"yy":1.0,"dx":0.0,"dy":0.0},"brush":{"LinearGradient":{"p0":{"x":500.0,"y":250.0},"p1":{"x":500.0,"y":950.0},"color_stops":[{"offset":0.0,"palette_index":0,"alpha":1.0},{"offset":1.0,"palette_index":4,"alpha":1.0}],"extend":"Repeat"}}}},"PopTransform","PopClip"]}
diff --git a/font-test-data/test_data/colrv1_json/colrv1_variable_alpha_APH2_-0.7_APH3_-0.2 b/font-test-data/test_data/colrv1_json/colrv1_variable_alpha_APH2_-0.7_APH3_-0.2
index 734cdfcc3..4ac73f701 100644
--- a/font-test-data/test_data/colrv1_json/colrv1_variable_alpha_APH2_-0.7_APH3_-0.2
+++ b/font-test-data/test_data/colrv1_json/colrv1_variable_alpha_APH2_-0.7_APH3_-0.2
@@ -1 +1 @@
-{"glyph_id":177,"ops":[{"PushClipBox":{"clip_box":{"x_min":0.0,"y_min":0.0,"x_max":1000.0,"y_max":1000.0}}},{"PushTransform":{"transform":{"xx":1.0,"yx":0.0,"xy":0.0,"yy":1.0,"dx":150.0,"dy":0.0}}},{"PushClipGlyph":{"gid":176}},{"FillBrush":{"brush":{"Solid":{"palette_index":3,"alpha":1.0}}}},"PopClip","PopTransform",{"PushTransform":{"transform":{"xx":1.0,"yx":0.0,"xy":0.0,"yy":1.0,"dx":-150.0,"dy":0.0}}},{"PushClipGlyph":{"gid":176}},{"FillBrush":{"brush":{"LinearGradient":{"p0":{"x":500.0,"y":250.0},"p1":{"x":500.0,"y":950.0},"color_stops":[{"offset":0.0,"palette_index":0,"alpha":0.2999878},{"offset":1.0,"palette_index":4,"alpha":0.7999878}],"extend":"Repeat"}}}},"PopClip","PopTransform","PopClip"]}
+{"glyph_id":177,"ops":[{"PushClipBox":{"clip_box":{"x_min":0.0,"y_min":0.0,"x_max":1000.0,"y_max":1000.0}}},{"PushTransform":{"transform":{"xx":1.0,"yx":0.0,"xy":0.0,"yy":1.0,"dx":150.0,"dy":0.0}}},{"FillGlyph":{"gid":176,"transform":{"xx":1.0,"yx":0.0,"xy":0.0,"yy":1.0,"dx":0.0,"dy":0.0},"brush":{"Solid":{"palette_index":3,"alpha":1.0}}}},"PopTransform",{"PushTransform":{"transform":{"xx":1.0,"yx":0.0,"xy":0.0,"yy":1.0,"dx":-150.0,"dy":0.0}}},{"FillGlyph":{"gid":176,"transform":{"xx":1.0,"yx":0.0,"xy":0.0,"yy":1.0,"dx":0.0,"dy":0.0},"brush":{"LinearGradient":{"p0":{"x":500.0,"y":250.0},"p1":{"x":500.0,"y":950.0},"color_stops":[{"offset":0.0,"palette_index":0,"alpha":0.2999878},{"offset":1.0,"palette_index":4,"alpha":0.7999878}],"extend":"Repeat"}}}},"PopTransform","PopClip"]}
diff --git a/skrifa/src/color/mod.rs b/skrifa/src/color/mod.rs
index 7c1441370..01805600d 100644
--- a/skrifa/src/color/mod.rs
+++ b/skrifa/src/color/mod.rs
@@ -218,7 +218,34 @@ pub trait ColorPainter {
     fn pop_clip(&mut self);
 
     /// Fill the current clip area with the specified gradient fill.
-    fn fill(&mut self, brush: Brush);
+    fn fill(&mut self, brush: Brush<'_>);
+
+    /// Combined clip and fill operation.
+    ///
+    /// Apply the clip path determined by the specified `glyph_id`, then fill it
+    /// with the specified [`brush`](Brush), applying the `_brush_transform`
+    /// transformation matrix to the brush. The default implementation works
+    /// based on existing methods in this trait. It is recommended for clients
+    /// to override the default implementaition with a custom combined clip and
+    /// fill operation. In this way overriding likely results in performance
+    /// gains depending on performance characteristics of the 2D graphics stack
+    /// that these calls are mapped to.
+    fn fill_glyph(
+        &mut self,
+        glyph_id: GlyphId,
+        brush_transform: Option<Transform>,
+        brush: Brush<'_>,
+    ) {
+        self.push_clip_glyph(glyph_id);
+        if let Some(wrap_in_transform) = brush_transform {
+            self.push_transform(wrap_in_transform);
+            self.fill(brush);
+            self.pop_transform();
+        } else {
+            self.fill(brush);
+        }
+        self.pop_clip();
+    }
 
     /// Optionally implement this method: Draw an unscaled COLRv1 glyph given
     /// the current transformation matrix (as accumulated by
@@ -230,9 +257,6 @@ pub trait ColorPainter {
         Ok(PaintCachedColorGlyph::Unimplemented)
     }
 
-    // TODO(https://github.com/googlefonts/fontations/issues/746):
-    // Add an optimized callback function combining clip, fill and transforms.
-
     /// Open a new layer, and merge the layer down using `composite_mode` when
     /// [`pop_layer`](ColorPainter::pop_layer) is called, signalling that this layer is done drawing.
     fn push_layer(&mut self, composite_mode: CompositeMode);
diff --git a/skrifa/src/color/transform.rs b/skrifa/src/color/transform.rs
index 8c55c0a8c..78bd1ef6b 100644
--- a/skrifa/src/color/transform.rs
+++ b/skrifa/src/color/transform.rs
@@ -1,4 +1,6 @@
 //! Contains a [`Transform`] object holding values of an affine transformation matrix.
+use std::ops::{Mul, MulAssign};
+
 use read_fonts::ReadError;
 
 use super::instance::ResolvedPaint;
@@ -8,7 +10,7 @@ use serde::{Deserialize, Serialize};
 
 #[derive(Clone, Debug, PartialEq)]
 #[cfg_attr(test, derive(Serialize, Deserialize))]
-/// A transformation matrix to be applied to the drawing canvas.  
+/// A transformation matrix to be applied to the drawing canvas.
 ///
 /// Factors are specified in column-order, meaning that
 /// for a vector `(x,y)` the transformed position `x'` of the vector
@@ -16,6 +18,7 @@ use serde::{Deserialize, Serialize};
 /// `x' = xx * x + xy * y + dx`,
 /// and the transformed position y' is calculated by
 /// `y' = yx * x + yy * y + dy`.
+#[derive(Copy)]
 pub struct Transform {
     pub xx: f32,
     pub yx: f32,
@@ -25,6 +28,30 @@ pub struct Transform {
     pub dy: f32,
 }
 
+impl MulAssign for Transform {
+    fn mul_assign(&mut self, rhs: Self) {
+        *self = *self * rhs;
+    }
+}
+
+impl Mul for Transform {
+    type Output = Self;
+
+    fn mul(self, rhs: Self) -> Self::Output {
+        fn muladdmul(a: f32, b: f32, c: f32, d: f32) -> f32 {
+            a * b + c * d
+        }
+        Self {
+            xx: muladdmul(self.xx, rhs.xx, self.xy, rhs.yx),
+            xy: muladdmul(self.xx, rhs.xy, self.xy, rhs.yy),
+            dx: muladdmul(self.xx, rhs.dx, self.xy, rhs.dy) + self.dx,
+            yx: muladdmul(self.yx, rhs.xx, self.yy, rhs.yx),
+            yy: muladdmul(self.yx, rhs.xy, self.yy, rhs.yy),
+            dy: muladdmul(self.yx, rhs.dx, self.yy, rhs.dy) + self.dy,
+        }
+    }
+}
+
 impl Default for Transform {
     fn default() -> Self {
         Transform {
diff --git a/skrifa/src/color/traversal.rs b/skrifa/src/color/traversal.rs
index 86bd81ace..8149b097a 100644
--- a/skrifa/src/color/traversal.rs
+++ b/skrifa/src/color/traversal.rs
@@ -13,7 +13,7 @@ use super::{
     instance::{
         resolve_clip_box, resolve_paint, ColorStops, ColrInstance, ResolvedColorStop, ResolvedPaint,
     },
-    Brush, ColorPainter, ColorStop, PaintCachedColorGlyph, PaintError,
+    Brush, ColorPainter, ColorStop, PaintCachedColorGlyph, PaintError, Transform,
 };
 
 // Workaround for https://bugs.chromium.org/p/chromium/issues/detail?id=1516634.
@@ -52,6 +52,73 @@ fn make_sorted_resolved_stops(stops: &ColorStops, instance: &ColrInstance) -> Ve
     collected
 }
 
+struct CollectFillGlyphPainter<'a> {
+    brush_transform: Option<Transform>,
+    glyph_id: GlyphId,
+    parent_painter: &'a mut dyn ColorPainter,
+    pub optimization_success: bool,
+}
+
+impl<'a> CollectFillGlyphPainter<'a> {
+    fn new(parent_painter: &'a mut dyn ColorPainter, glyph_id: GlyphId) -> Self {
+        Self {
+            brush_transform: None,
+            glyph_id,
+            parent_painter,
+            optimization_success: true,
+        }
+    }
+}
+
+impl<'a> ColorPainter for CollectFillGlyphPainter<'a> {
+    fn push_transform(&mut self, transform: Transform) {
+        if self.optimization_success {
+            match self.brush_transform {
+                None => {
+                    self.brush_transform = Some(transform);
+                }
+                Some(ref mut existing_transform) => {
+                    *existing_transform *= transform;
+                }
+            }
+        }
+    }
+
+    fn pop_transform(&mut self) {
+        // Since we only support fill and and transform operations, we need to
+        // ignore a popped transform, as this would be called after traversing
+        // the graph backup after a fill was performed, but we want to preserve
+        // the transform in order to be able to return it.
+    }
+
+    fn fill(&mut self, brush: Brush<'_>) {
+        if self.optimization_success {
+            self.parent_painter
+                .fill_glyph(self.glyph_id, self.brush_transform.clone(), brush);
+        }
+    }
+
+    fn push_clip_glyph(&mut self, _: GlyphId) {
+        self.optimization_success = false;
+    }
+
+    fn push_clip_box(&mut self, _: BoundingBox<f32>) {
+        self.optimization_success = false;
+    }
+
+    fn pop_clip(&mut self) {
+        self.optimization_success = false;
+    }
+
+    fn push_layer(&mut self, _: CompositeMode) {
+        self.optimization_success = false;
+    }
+
+    fn pop_layer(&mut self) {
+        self.optimization_success = false;
+    }
+}
+
 pub(crate) fn traverse_with_callbacks(
     paint: &ResolvedPaint,
     instance: &ColrInstance,
@@ -365,14 +432,26 @@ pub(crate) fn traverse_with_callbacks(
         }
 
         ResolvedPaint::Glyph { glyph_id, paint } => {
-            painter.push_clip_glyph(*glyph_id);
-            let result = traverse_with_callbacks(
+            let mut optimizer = CollectFillGlyphPainter::new(painter, *glyph_id);
+            let mut result = traverse_with_callbacks(
                 &resolve_paint(instance, paint)?,
                 instance,
-                painter,
+                &mut optimizer,
                 visited_set,
             );
-            painter.pop_clip();
+
+            // In case the optimization was not successful, just push a clip, and continue unoptimized traversal.
+            if !optimizer.optimization_success {
+                painter.push_clip_glyph(*glyph_id);
+                result = traverse_with_callbacks(
+                    &resolve_paint(instance, paint)?,
+                    instance,
+                    painter,
+                    visited_set,
+                );
+                painter.pop_clip();
+            }
+
             result
         }
         ResolvedPaint::ColrGlyph { glyph_id } => match (*instance).v1_base_glyph(*glyph_id)? {
@@ -467,14 +546,14 @@ pub(crate) fn traverse_v0_range(
 ) -> Result<(), PaintError> {
     for layer_index in range.clone() {
         let (layer_index, palette_index) = (*instance).v0_layer(layer_index)?;
-        // TODO(https://github.com/googlefonts/fontations/issues/746):
-        // Use optimized callback function combining clip, fill and transforms.
-        painter.push_clip_glyph(layer_index);
-        painter.fill(Brush::Solid {
-            palette_index,
-            alpha: 1.0,
-        });
-        painter.pop_clip();
+        painter.fill_glyph(
+            layer_index,
+            None,
+            Brush::Solid {
+                palette_index,
+                alpha: 1.0,
+            },
+        );
     }
     Ok(())
 }
diff --git a/skrifa/src/color/traversal_tests/mod.rs b/skrifa/src/color/traversal_tests/mod.rs
index 12b21ea74..71d45fe53 100644
--- a/skrifa/src/color/traversal_tests/mod.rs
+++ b/skrifa/src/color/traversal_tests/mod.rs
@@ -147,13 +147,28 @@ impl From<Transform> for DumpTransform {
 
 #[derive(Serialize, Deserialize, PartialEq)]
 enum PaintOps {
-    PushTransform { transform: DumpTransform },
+    PushTransform {
+        transform: DumpTransform,
+    },
     PopTransform,
-    PushClipGlyph { gid: u16 },
-    PushClipBox { clip_box: BoundingBox<f32> },
+    PushClipGlyph {
+        gid: u16,
+    },
+    PushClipBox {
+        clip_box: BoundingBox<f32>,
+    },
     PopClip,
-    FillBrush { brush: BrushParams },
-    PushLayer { composite_mode: u8 },
+    FillBrush {
+        brush: BrushParams,
+    },
+    FillGlyph {
+        gid: u16,
+        transform: DumpTransform,
+        brush: BrushParams,
+    },
+    PushLayer {
+        composite_mode: u8,
+    },
     PopLayer,
 }
 
@@ -187,6 +202,14 @@ impl ColorPainter for PaintDump {
         });
     }
 
+    fn fill_glyph(&mut self, glyph_id: GlyphId, transform: Option<Transform>, brush: Brush) {
+        self.ops.push(PaintOps::FillGlyph {
+            gid: glyph_id.to_u16(),
+            transform: transform.unwrap_or_default().into(),
+            brush: brush.into(),
+        });
+    }
+
     fn push_layer(&mut self, composite_mode: CompositeMode) {
         self.ops.push(PaintOps::PushLayer {
             composite_mode: composite_mode as u8,