diff --git a/pytype/pyi/definitions.py b/pytype/pyi/definitions.py
index d6922a3cc..7c5a9d83b 100644
--- a/pytype/pyi/definitions.py
+++ b/pytype/pyi/definitions.py
@@ -468,11 +468,13 @@ def add_type_variable(self, name, tvar):
       raise _ParseError(f"{tvar.kind} name needs to be {tvar.name!r} "
                         f"(not {name!r})")
     bound = tvar.bound
-    if isinstance(bound, str):
-      bound = pytd.NamedType(bound)
     constraints = tuple(tvar.constraints) if tvar.constraints else ()
+    if isinstance(tvar.default, list):
+      default = tuple(tvar.default)
+    else:
+      default = tvar.default
     self.type_params.append(pytd_type(
-        name=name, constraints=constraints, bound=bound))
+        name=name, constraints=constraints, bound=bound, default=default))
 
   def add_import(self, from_package, import_list):
     """Add an import.
diff --git a/pytype/pyi/parser.py b/pytype/pyi/parser.py
index 89ff9a8ed..c1af66698 100644
--- a/pytype/pyi/parser.py
+++ b/pytype/pyi/parser.py
@@ -8,7 +8,7 @@
 import re
 import sys
 import tokenize
-from typing import Any, List, Optional, Tuple, cast
+from typing import Any, List, Optional, Tuple, Union, cast
 
 from pytype.ast import debug
 from pytype.pyi import conditions
@@ -61,6 +61,7 @@ class _TypeVariable:
   name: str
   bound: Optional[pytd.Type]
   constraints: List[pytd.Type]
+  default: Optional[Union[pytd.Type, List[pytd.Type]]]
 
   @classmethod
   def from_call(cls, kind: str, node: astlib.Call):
@@ -72,18 +73,20 @@ def from_call(cls, kind: str, node: astlib.Call):
     if not types.Pyval.is_str(name):
       raise ParseError(f"Bad arguments to {kind}")
     bound = None
-    # 'bound' is the only keyword argument we currently use.
+    default = None
     # TODO(rechen): We should enforce the PEP 484 guideline that
     # len(constraints) != 1. However, this guideline is currently violated
     # in typeshed (see https://github.com/python/typeshed/pull/806).
     kws = {x.arg for x in node.keywords}
-    extra = kws - {"bound", "covariant", "contravariant"}
+    extra = kws - {"bound", "covariant", "contravariant", "default"}
     if extra:
       raise ParseError(f"Unrecognized keyword(s): {', '.join(extra)}")
     for kw in node.keywords:
       if kw.arg == "bound":
         bound = kw.value
-    return cls(kind, name.value, bound, constraints)
+      elif kw.arg == "default":
+        default = kw.value
+    return cls(kind, name.value, bound, constraints, default)
 
 #------------------------------------------------------
 # Main tree visitor and generator code
@@ -674,6 +677,8 @@ def _convert_typevar_args(self, node: astlib.Call):
     for kw in node.keywords:
       if kw.arg == "bound":
         kw.value = self.annotation_visitor.visit(kw.value)
+      elif kw.arg == "default":
+        kw.value = self.annotation_visitor.visit(kw.value)
 
   def _convert_typed_dict_args(self, node: astlib.Call):
     for fields in node.args[1:]:
@@ -682,7 +687,8 @@ def _convert_typed_dict_args(self, node: astlib.Call):
   def enter_Call(self, node):
     node.func = self.annotation_visitor.visit(node.func)
     func = node.func.name or ""
-    if self.defs.matches_type(func, ("typing.TypeVar", "typing.ParamSpec")):
+    if self.defs.matches_type(func, ("typing.TypeVar", "typing.ParamSpec",
+                                     "typing.TypeVarTuple")):
       self._convert_typevar_args(node)
     elif self.defs.matches_type(func, "typing.NamedTuple"):
       self._convert_typing_namedtuple_args(node)
diff --git a/pytype/pyi/parser_test.py b/pytype/pyi/parser_test.py
index 1cd278aad..bd3aa2056 100644
--- a/pytype/pyi/parser_test.py
+++ b/pytype/pyi/parser_test.py
@@ -3526,5 +3526,30 @@ def f(x: Tuple[Any, ...]) -> Any: ...
     """)
 
 
+class TypeParameterDefaultTest(parser_test_base.ParserTestBase):
+
+  def test_typevar(self):
+    self.check("""
+      from typing_extensions import TypeVar
+
+      T = TypeVar('T', default=int)
+    """)
+
+  def test_paramspec(self):
+    self.check("""
+      from typing_extensions import ParamSpec
+
+      P = ParamSpec('P', default=[str, int])
+    """)
+
+  def test_typevartuple(self):
+    self.check("""
+      from typing_extensions import TypeVarTuple, Unpack
+      Ts = TypeVarTuple('Ts', default=Unpack[tuple[str, int]])
+    """, """
+      from typing_extensions import TypeVarTuple, TypeVarTuple as Ts, Unpack
+    """)
+
+
 if __name__ == "__main__":
   unittest.main()
diff --git a/pytype/pytd/printer.py b/pytype/pytd/printer.py
index 9ae082226..226255c74 100644
--- a/pytype/pytd/printer.py
+++ b/pytype/pytd/printer.py
@@ -209,6 +209,11 @@ def _FormatTypeParams(self, type_params):
       args += [self.Print(c) for c in t.constraints]
       if t.bound:
         args.append(f"bound={self.Print(t.bound)}")
+      if isinstance(t.default, tuple):
+        args.append(
+            f"default=[{', '.join(self.Print(d) for d in t.default)}]")
+      elif t.default:
+        args.append(f"default={self.Print(t.default)}")
       if isinstance(t, pytd.ParamSpec):
         typename = self._LookupTypingMember("ParamSpec")
       else:
diff --git a/pytype/pytd/pytd.py b/pytype/pytd/pytd.py
index 56f505e77..3a6b4a33a 100644
--- a/pytype/pytd/pytd.py
+++ b/pytype/pytd/pytd.py
@@ -344,6 +344,7 @@ def f(x: T) -> T
   name: str
   constraints: Tuple[TypeU, ...] = ()
   bound: Optional[TypeU] = None
+  default: Optional[Union[TypeU, Tuple[TypeU, ...]]] = None
   scope: Optional[str] = None
 
   def __lt__(self, other):
diff --git a/pytype/pytd/visitors_test.py b/pytype/pytd/visitors_test.py
index 28bcd4846..576ca2791 100644
--- a/pytype/pytd/visitors_test.py
+++ b/pytype/pytd/visitors_test.py
@@ -552,8 +552,7 @@ class A(Dict[T, T], Generic[T]): pass
     """)
     a = ast.Lookup("A")
     self.assertEqual(
-        (pytd.TemplateItem(pytd.TypeParameter("T", (), None, "A")),),
-        a.template)
+        (pytd.TemplateItem(pytd.TypeParameter("T", scope="A")),), a.template)
 
   def test_adjust_type_parameters_with_duplicates_in_generic(self):
     src = textwrap.dedent("""